Instituto de Química del Noroeste Argentino

Abstract Observations show that glaciers around the world are in retreat and losing mass. Internationally coordinated for over a century, glacier monitoring activities provide an unprecedented dataset of glacier observations from ground, air and space. Glacier studies generally select specific parts of these datasets to obtain optimal assessments of the mass-balance data relating to the impact that glaciers exercise on global sea-level fluctuations or on regional runoff. In this study we provide an overview and analysis of the main observational datasets compiled by the World Glacier Monitoring Service (WGMS). The dataset on glacier front variations (∼42 000 since 1600) delivers clear evidence that centennial glacier retreat is a global phenomenon. Intermittent readvance periods at regional and decadal scale are normally restricted to a subsample of glaciers and have not come close to achieving the maximum positions of the Little Ice Age (or Holocene). Glaciological and geodetic observations (∼5200 since 1850) show that the rates of early 21st-century mass loss are without precedent on a global scale, at least for the time period observed and probably also for recorded history, as indicated also in reconstructions from written and illustrated documents. This strong imbalance implies that glaciers in many regions will very likely suffer further ice loss, even if climate remains stable.

Severe droughts have the potential to reduce forest productivity and trigger tree mortality. Most trees face several drought events during their life and therefore resilience to dry conditions may be crucial to long-term survival. We assessed how growth resilience to severe droughts, including its components resistance and recovery, is related to the ability to survive future droughts by using a tree-ring database of surviving and now-dead trees from 118 sites (22 species, >3,500 trees). We found that, across the variety of regions and species sampled, trees that died during water shortages were less resilient to previous non-lethal droughts, relative to coexisting surviving trees of the same species. In angiosperms, drought-related mortality risk is associated with lower resistance (low capacity to reduce impact of the initial drought), while it is related to reduced recovery (low capacity to attain pre-drought growth rates) in gymnosperms. The different resilience strategies in these two taxonomic groups open new avenues to improve our understanding and prediction of drought-induced mortality.

• The advent of embryophytes (land plants) is among the most important evolutionary breakthroughs in Earth history. It irreversibly changed climates and biogeochemical processes on a global scale; it allowed all eukaryotic terrestrial life to evolve and to invade nearly all continental environments. Before this work, the earliest unequivocal embryophyte traces were late Darriwilian (late Middle Ordovician; c. 463-461 million yr ago (Ma)) cryptospores from Saudi Arabia and from the Czech Republic (western Gondwana). • Here, we processed Dapingian (early Middle Ordovician, c. 473-471 Ma) palynological samples from Argentina (eastern Gondwana). • We discovered a diverse cryptospore assemblage, including naked and envelope-enclosed monads and tetrads, representing five genera. • Our discovery reinforces the earlier suggestion that embryophytes first evolved in Gondwana. It indicates that the terrestrialization of plants might have begun in the eastern part of Gondwana. The diversity of the Dapingian assemblage implies an earlier, Early Ordovician or even Cambrian, origin of embryophytes. Dapingian to Aeronian (Early Silurian) cryptospore assemblages are similar, suggesting that the rate of embryophyte evolution was extremely slow during the first c. 35-45 million yr of their diversification. The Argentinean cryptospores predate other cryptospore occurrences by c. 8-12 million yr, and are currently the earliest evidence of plants on land.

Abstract Atmospheric rivers (ARs) are now widely known for their association with high‐impact weather events and long‐term water supply in many regions. Researchers within the scientific community have developed numerous methods to identify and track of ARs—a necessary step for analyses on gridded data sets, and objective attribution of impacts to ARs. These different methods have been developed to answer specific research questions and hence use different criteria (e.g., geometry, threshold values of key variables, and time dependence). Furthermore, these methods are often employed using different reanalysis data sets, time periods, and regions of interest. The goal of the Atmospheric River Tracking Method Intercomparison Project (ARTMIP) is to understand and quantify uncertainties in AR science that arise due to differences in these methods. This paper presents results for key AR‐related metrics based on 20+ different AR identification and tracking methods applied to Modern‐Era Retrospective Analysis for Research and Applications Version 2 reanalysis data from January 1980 through June 2017. We show that AR frequency, duration, and seasonality exhibit a wide range of results, while the meridional distribution of these metrics along selected coastal (but not interior) transects are quite similar across methods. Furthermore, methods are grouped into criteria‐based clusters, within which the range of results is reduced. AR case studies and an evaluation of individual method deviation from an all‐method mean highlight advantages/disadvantages of certain approaches. For example, methods with less (more) restrictive criteria identify more (less) ARs and AR‐related impacts. Finally, this paper concludes with a discussion and recommendations for those conducting AR‐related research to consider.

, possibly due to tree growth stimulation. Extant models predict that this growth stimulation will continue to cause a net carbon uptake this century. However, there are indications that increased growth rates may shorten trees' lifespan and thus recent increases in forest carbon stocks may be transient due to lagged increases in mortality. Here we show that growth-lifespan trade-offs are indeed near universal, occurring across almost all species and climates. This trade-off is directly linked to faster growth reducing tree lifespan, and not due to covariance with climate or environment. Thus, current tree growth stimulation will, inevitably, result in a lagged increase in canopy tree mortality, as is indeed widely observed, and eventually neutralise carbon gains due to growth stimulation. Results from a strongly data-based forest simulator confirm these expectations. Extant Earth system model projections of global forest carbon sink persistence are likely too optimistic, increasing the need to curb greenhouse gas emissions.

Abstract The snowpack in the central Andes (30°–37°S) is the primary source for streamflow in central Chile and central-western Argentina, but few published studies are available on snowpack variability in the region. This paper develops the first regional snowpack series (1951–2005) from Chilean and Argentinean snow course records. This series shows a strong regional signal, marked interannual variability, and a positive, though nonsignificant, linear trend. Correlations with local precipitation and temperature records reveal a marked association with conditions in central Chile. High snow accumulation is generally concurrent with El Niño events in the tropical Pacific, but only 5 of the 10 driest years coincided with La Niña events. Evaluation of 500-hPa geopotential height anomaly maps during extreme snow years highlights the crucial significance of tropospheric conditions in the subtropical and southeast Pacific in modulating snowfall. Correlations with gridded SST and SLP data and multiple regressions with large-scale climatic indices corroborate a Pacific ENSO-related influence largely concentrated during the austral winter months. This hampers the predictability of snowpack before the onset of the cold season. Annual and warm-season river discharges on both sides of the cordillera are significantly correlated with the regional snowpack record and show positive linear trends over the 1951–2004 common period, probably related to a greater frequency of above-average snowpacks during recent decades. Future demand and competition for water resources in these highly populated regions will require detailed information about temporal and spatial variations in snow accumulation over the Andes. The results indicate that the relationships between snowpack and atmospheric circulation patterns prior to the winter season are complex, and more detailed analyses are necessary to improve prediction of winter snowfall totals.

Propolis is one of the most fascinating honey bee (Apis mellifera L.) products. It is a plant derived product that bees produce from resins that they collect from different plant organs and with which they mix beeswax. Propolis is a building material and a protective agent in the bee hive. It also plays an important role in honey bee social immunity, and is widely used by humans as an ingredient of nutraceuticals, over-the-counter preparations and cosmetics. Its chemical
\ncomposition varies by geographic location, climatic zone and local flora. The understanding of the chemical diversity of propolis is very important in propolis research. In this manuscript, we give an overview of the available methods for studying propolis in different aspects: propolis in the bee colony; chemical composition and plant sources of propolis; biological activity of propolis with respect to bees and humans; and approaches for standardization and quality control for the purposes of industrial application.

South American (SA) societies are highly vulnerable to droughts and pluvials, but lack of long-term climate observations severely limits our understanding of the global processes driving climatic variability in the region. The number and quality of SA climate-sensitive tree ring chronologies have significantly increased in recent decades, now providing a robust network of 286 records for characterizing hydroclimate variability since 1400 CE. We combine this network with a self-calibrated Palmer Drought Severity Index (scPDSI) dataset to derive the South American Drought Atlas (SADA) over the continent south of 12°S. The gridded annual reconstruction of austral summer scPDSI is the most spatially complete estimate of SA hydroclimate to date, and well matches past historical dry/wet events. Relating the SADA to the Australia-New Zealand Drought Atlas, sea surface temperatures and atmospheric pressure fields, we determine that the El Niño-Southern Oscillation (ENSO) and the Southern Annular Mode (SAM) are strongly associated with spatially extended droughts and pluvials over the SADA domain during the past several centuries. SADA also exhibits more extended severe droughts and extreme pluvials since the mid-20th century. Extensive droughts are consistent with the observed 20th-century trend toward positive SAM anomalies concomitant with the weakening of midlatitude Westerlies, while low-level moisture transport intensified by global warming has favored extreme rainfall across the subtropics. The SADA thus provides a long-term context for observed hydroclimatic changes and for 21st-century Intergovernmental Panel on Climate Change (IPCC) projections that suggest SA will experience more frequent/severe droughts and rainfall events as a consequence of increasing greenhouse gas emissions.

Abstract. The detachment of large parts of low-angle mountain glaciers resulting in massive ice–rock avalanches have so far been believed to be a unique type of event, made known to the global scientific community first for the 2002 Kolka Glacier detachment, Caucasus Mountains, and then for the 2016 collapses of two glaciers in the Aru range, Tibet. Since 2016, several so-far unrecognized low-angle glacier detachments have been recognized and described, and new ones have occurred. In the current contribution, we compile, compare, and discuss 20 actual or suspected large-volume detachments of low-angle mountain glaciers at 10 different sites in the Caucasus, the Pamirs, Tibet, Altai, the North American Cordillera, and the Southern Andes. Many of the detachments reached volumes in the order of 10–100 million m3. The similarities and differences between the presented cases indicate that glacier detachments often involve a coincidental combination of factors related to the lowering of basal friction, high or increasing driving stresses, concentration of shear stress, or low resistance to exceed stability thresholds. Particularly soft glacier beds seem to be a common condition among the observed events as they offer smooth contact areas between the glacier and the underlying substrate and are prone to till-strength weakening and eventually basal failure under high pore-water pressure. Partially or fully thawed glacier bed conditions and the presence of liquid water could thus play an important role in the detachments. Surface slopes of the detached glaciers range between around 10∘ and 20∘. This may be low enough to enable the development of thick and thus large-volume glaciers while also being steep enough to allow critical driving stresses to build up. We construct a simple slab model to estimate ranges of glacier slope and width above which a glacier may be able to detach when extensively losing basal resistance. From this model we estimate that all the detachments described in this study occurred due to a basal shear stress reduction of more than 50 %. Most of the ice–rock avalanches resulting from the detachments in this study have a particularly low angle of reach, down to around 5∘, likely due to their high ice content and connected liquefaction potential, the availability of soft basal slurries, and large amounts of basal water, as well as the smooth topographic setting typical for glacial valleys. Low-angle glacier detachments combine elements and likely also physical processes of glacier surges and ice break-offs from steep glaciers. The surge-like temporal evolution ahead of several detachments and their geographic proximity to other surge-type glaciers indicate the glacier detachments investigated can be interpreted as endmembers of the continuum of surge-like glacier instabilities. Though rare, glacier detachments appear to be more frequent than commonly thought and disclose, despite local differences in conditions and precursory evolutions, the fundamental and critical potential of low-angle soft glacier beds to fail catastrophically.

The Andes is the longest cordillera in the world and extends from northern South America (11°N) to the southern tip of the continent (53°S). The Andes runs through seven countries and is characterized by a wide variety of ecosystems strongly related to the contrasting climate over its eastern and western sides and along its latitudinal extension. In fact, the tropical Andes is the most biodiverse region on Earth. Currently, this region faces the highest potential impact of climate change, which could affect food security and water supplies for about 90 million people. From a scientific and societal view, the Andes present specific challenges because of its unique landscape and the fragile equilibrium between the growing population and its environment. In this manuscript, we provide an updated review of the most relevant scientific literature regarding the hydroclimate of the Andes. This review paper is presented in two parts. Part I is dedicated to summarize the scientific knowledge about the main climatic features of the Andes, with emphasis on mean large-scale atmospheric circulation, the Andes-Amazon hydroclimate interconnections, and the regular cycles of precipitation, including the most characteristic diurnal and annual cycles of precipitation. Part II, which is also included in the research topic “Connecting Mountain Hydroclimate Through the American Cordilleras”, focuses on hydroclimate variability of the Andes at a sub-continental scale.

Abstract This study quantifies the impact of atmospheric rivers (ARs) on precipitation in southern South America. An AR detection algorithm was developed based on integrated water vapor transport (IVT) from 6-hourly CFSR reanalysis data over a 16-yr period (2001–16). AR landfalls were linked to precipitation using a comprehensive observing network that spanned large variations in terrain along and across the Andes from 27° to 55°S, including some sites with hourly data. Along the Pacific (west) coast, AR landfalls are most frequent between 38° and 50°S, averaging 35–40 days yr−1. This decreases rapidly to the south and north of this maximum, as well as to the east of the Andes. Landfalling ARs are more frequent in winter/spring (summer/fall) to the north (south) of ~43°S. ARs contribute 45%–60% of the annual precipitation in subtropical Chile (37°–32°S) and 40%–55% along the midlatitude west coast (37°–47°S). These values significantly exceed those in western North America, likely due to the Andes being taller. In subtropical and midlatitude regions, roughly half of all events with top-quartile precipitation rates occur under AR conditions. Median daily and hourly precipitation in ARs is 2–3 times that of other storms. The results of this study extend knowledge of the key roles of ARs on precipitation, weather, and climate in the South American region. They enable comparisons with other areas globally, provide context for specific events, and support local nowcasting and forecasting.

This paper describes spatial distribution patterns of the phytoplankton community (composition, cell abundance and biomass concentration) in relation to local environmental conditions in the Southern Ocean. Sampling was performed during summer 1997 off the coast of the western Antarctic Peninsula between Anvers Island and Marguerite Bay. Phytoplankton was characterized by relatively low biomass throughout most of the study area and was dominated by nanoalgae (<20 μm). Phytoplankton varied along an on-offshore gradient, with decreasing total cell abundance, chlorophyll a (chl a) concentration and carbon biomass toward the open ocean. Chl a concentration showed surface or subsurface maxima in coastal and middle-shelf waters, and deep maxima between ∼40 and 100 m in oceanic waters. Across-shelf variability in phytoplankton correlated with vertical stability in the water column, which appears to be the major parameter affecting phytoplankton community structure in the area. We hypothesize that the deep chl a maximum offshore may be associated with iron limitation in near-surface waters and higher iron concentration in 'winter waters' (subsurface remnant of Antarctic Surface Waters). On a smaller spatial scale, a cluster analysis showed great regional variability in phytoplankton assemblages. The area was divided into 4 main regions based on differences in the phytoplankton composition and concentration. Three peaks in phytoplankton abundance were found on a north-to-south gradient in near-shore waters: a Cryptomonas spp. bloom near Anvers Island, a small unidentified phytoflagellate bloom in Grandidier Channel, and a diatom bloom in Marguerite Bay. These assemblages resemble different stages of the phytoplankton seasonal succession, and may be related to the progressive sea-ice retreat, which might have regulated the timing of the onset of the phytoplankton seasonal succession in a north-south gradient. Biological environmental factors, such as seeding of the water column by epontic algae and selective zooplankton herbivory, are hypothesized to affect community composition in coastal regions. We conclude that large-scale variability in phytoplankton community structure is related to water column physical conditions and possibly iron availability, while mesoscale variability, as seen in coastal waters, is more likely due to seasonal succession of different algae groups.

Abstract Though tree-ring chronologies are annually resolved, their dating has never been independently validated at the global scale. Moreover, it is unknown if atmospheric radiocarbon enrichment events of cosmogenic origin leave spatiotemporally consistent fingerprints. Here we measure the 14 C content in 484 individual tree rings formed in the periods 770–780 and 990–1000 CE. Distinct 14 C excursions starting in the boreal summer of 774 and the boreal spring of 993 ensure the precise dating of 44 tree-ring records from five continents. We also identify a meridional decline of 11-year mean atmospheric radiocarbon concentrations across both hemispheres. Corroborated by historical eye-witness accounts of red auroras, our results suggest a global exposure to strong solar proton radiation. To improve understanding of the return frequency and intensity of past cosmic events, which is particularly important for assessing the potential threat of space weather on our society, further annually resolved 14 C measurements are needed.

Abstract X‐ray microdensitometry on annually resolved tree‐ring samples has gained an exceptional position in last‐millennium paleoclimatology through the maximum latewood density (MXD) parameter, but also increasingly through other density parameters. For 50 years, X‐ray based measurement techniques have been the de facto standard. However, studies report offsets in the mean levels for MXD measurements derived from different laboratories, indicating challenges of accuracy and precision. Moreover, reflected visible light‐based techniques are becoming increasingly popular, and wood anatomical techniques are emerging as a potentially powerful pathway to extract density information at the highest resolution. Here we review the current understanding and merits of wood density for tree‐ring research, associated microdensitometric techniques, and analytical measurement challenges. The review is further complemented with a careful comparison of new measurements derived at 17 laboratories, using several different techniques. The new experiment allowed us to corroborate and refresh “long‐standing wisdom” but also provide new insights. Key outcomes include (i) a demonstration of the need for mass/volume‐based recalibration to accurately estimate average ring density; (ii) a substantiation of systematic differences in MXD measurements that cautions for great care when combining density data sets for climate reconstructions; and (iii) insights into the relevance of analytical measurement resolution in signals derived from tree‐ring density data. Finally, we provide recommendations expected to facilitate futureinter‐comparability and interpretations for global change research.

Abstract. The seasonal climate drivers of the carbon cycle in tropical forests remain poorly known, although these forests account for more carbon assimilation and storage than any other terrestrial ecosystem. Based on a unique combination of seasonal pan-tropical data sets from 89 experimental sites (68 include aboveground wood productivity measurements and 35 litter productivity measurements), their associated canopy photosynthetic capacity (enhanced vegetation index, EVI) and climate, we ask how carbon assimilation and aboveground allocation are related to climate seasonality in tropical forests and how they interact in the seasonal carbon cycle. We found that canopy photosynthetic capacity seasonality responds positively to precipitation when rainfall is < 2000 mm yr−1 (water-limited forests) and to radiation otherwise (light-limited forests). On the other hand, independent of climate limitations, wood productivity and litterfall are driven by seasonal variation in precipitation and evapotranspiration, respectively. Consequently, light-limited forests present an asynchronism between canopy photosynthetic capacity and wood productivity. First-order control by precipitation likely indicates a decrease in tropical forest productivity in a drier climate in water-limited forest, and in current light-limited forest with future rainfall < 2000 mm yr−1.

KEY MESSAGE: This review provides a comprehensive atlas of QTLs, genes, and alleles conferring resistance to 28 important diseases in all major soybean production regions in the world. Breeding disease-resistant soybean [Glycine max (L.) Merr.] varieties is a common goal for soybean breeding programs to ensure the sustainability and growth of soybean production worldwide. However, due to global climate change, soybean breeders are facing strong challenges to defeat diseases. Marker-assisted selection and genomic selection have been demonstrated to be successful methods in quickly integrating vertical resistance or horizontal resistance into improved soybean varieties, where vertical resistance refers to R genes and major effect QTLs, and horizontal resistance is a combination of major and minor effect genes or QTLs. This review summarized more than 800 resistant loci/alleles and their tightly linked markers for 28 soybean diseases worldwide, caused by nematodes, oomycetes, fungi, bacteria, and viruses. The major breakthroughs in the discovery of disease resistance gene atlas of soybean were also emphasized which include: (1) identification and characterization of vertical resistance genes reside rhg1 and Rhg4 for soybean cyst nematode, and exploration of the underlying regulation mechanisms through copy number variation and (2) map-based cloning and characterization of Rps11 conferring resistance to 80% isolates of Phytophthora sojae across the USA. In this review, we also highlight the validated QTLs in overlapping genomic regions from at least two studies and applied a consistent naming nomenclature for these QTLs. Our review provides a comprehensive summary of important resistant genes/QTLs and can be used as a toolbox for soybean improvement. Finally, the summarized genetic knowledge sheds light on future directions of accelerated soybean breeding and translational genomics studies.

In arid regions throughout the world, shallow phreatic aquifers feed natural oases of much higher productivity than would be expected solely from local rainfall. In South America, the presence of well-developed Prosopis flexuosa woodlands in the Monte Desert region east of the Andes has puzzled scientists for decades. Today these woodlands provide crucial subsistence to local populations, including descendants of the indigenous Huarpes. We explore the vulnerability and importance of phreatic groundwater for the productivity of the region, comparing the contributions of local rainfall to that of remote mountain recharge that is increasingly being diverted for irrigated agriculture before it reaches the desert. We combined deep soil coring, plant measurements, direct water-table observations, and stable-isotopic analyses (2H and 18O) of meteoric, surface, and ground waters at three study sites across the region, comparing woodland stands, bare dunes, and surrounding shrublands. The isotopic composition of phreatic groundwaters (delta2H: -137 per thousand +/- 5 per thousand) closely matched the signature of water brought to the region by the Mendoza River (-137 per thousand +/- 6 per thousand), suggestin that mountain-river infiltration rather than in situ rainfall deep drainage (-39 per thousand +/- 19 per thousand) was the dominant mechanism of recharge. Similarly, chloride mass balances determined from deep soil profiles (> 6 m) suggested very low recharge rates. Vegetation in woodland ecosystems, where significant groundwater discharge losses, likely >100 mm/yr occurred, relied on regionally derived groundwater located from 6.5 to 9.5 m underground. At these locations, daily water-table fluctuations of 10 mm, and stable-isotopic measurements of plant water, indicated groundwater uptake rates of 200-300 mm/yr. Regional scaling suggests that groundwater evapotranspiration reaches 18-42 mm/yr across the landscape, accounting for 7 17% of the Mendoza River flow regionally. Our study highlights the reliance of ecosystem productivity in natural oases on Andean snowmelt, which is increasingly being diverted to one of the largest irrigated regions of the continent. Understanding the ecohydrological coupling of mountain and desert ecosystems here and elsewhere should help managers balance production agriculture and conservation of unique woodland ecosystems and the rural communities that rely on them.

As a strategy to find efficient lignocellulose degrading enzymes/microorganisms for sugarcane biomass pretreatment purposes, 118 culturable bacterial strains were isolated from intestines of sugarcane-fed larvae of the moth Diatraea saccharalis. All strains were tested for cellulolytic activity using soluble carboxymethyl cellulose (CMC) degrading assays or by growing bacteria on sugarcane biomass as sole carbon sources. Out of the 118 strains isolated thirty eight were found to possess cellulose degrading activity and phylogenetic studies of the 16S rDNA sequence revealed that all cellulolytic strains belonged to the phyla γ-Proteobacteria, Actinobacteria and Firmicutes. Within the three phyla, species belonging to five different genera were identified (Klebsiella, Stenotrophomonas, Microbacterium, Bacillus and Enterococcus). Bacterial growth on sugarcane biomass as well as extracellular endo-glucanase activity induced on soluble cellulose was found to be highest in species belonging to genera Bacillus and Klebsiella. Good cellulolytic activity correlated with high extracellular protein concentrations. In addition, scanning microscopy studies revealed attachment of cellulolytic strains to different sugarcane substrates. The results of this study indicate the possibility to find efficient cellulose degrading enzymes and microorganisms from intestines of insect larvae feeding on sugarcane and their possible application in industrial processing of sugarcane biomass such as second generation biofuel production.

It is generally assumed that tree growth in the upper limit of a forest is mainly controlled by summer temperature. This general statement is mostly based on studies from extra-tropical mountains and has been rarely evaluated in subtropical latitudes frequently characterized by drier climates. In the subtropical mountains from Northwestern Argentina (∼23° S), annual precipitation decreases with elevation from >1500 mm at 1200– 1500 m, to <200 mm above 4000 m. In consequence, tree growth at high elevations in the region may be seriously limited by water supply. In order to assess the influence of precipitation on tree growth, we evaluated the relationships between climatic variations and radial growth in four species growing at different altitudinal zones: Juglans australis from the montane cloud forest at 1800 m; Alnus acuminata from the montane savanna-like woodland at 2700 m; Prosopis ferox from the subalpine dryland at 3500 m; and Polylepis tarapacana from the high-elevation alpine dryland at 4750 m. Dendrochronological techniques were used to relate variations in annual ring width with instrumental climatic records. Growth rings were correctly dated to the year of formation, and the cross-dated series standardized using autoregressive models to reduce non-climatic signals present in the records. Tree-ring chronologies, ranging from 117 to 341 years, were compared, during the common period, with instrumental climatic records using correlation-function analysis. In spite of the remarkable differences in elevation and environmental conditions among the sampling sites, correlation functions with climate indicated that the radial growth of the four species is largely controlled by precipitation. In most cases, increased precipitation during the previous and current growing seasons favors tree growth, while temperatures are negatively correlated with radial growth, likely due to the negative effect on water availability. These results indicate that the generalized idea of upper-treeline growth limited by summer temperatures should be carefully evaluated in low-latitude environments and does not apply to subtropical areas with severe water deficits or strong moisture seasonalities.

Abstract. Mendozasaurus neguyelap gen. et sp. nov. is a new titanosaur from the Upper Cretaceous of Neuquen Basin and the first known dinosaur species from Mendoza Province, Argentina. The remains were found in levels provisionally referred to the Rio Neuquen Formation (late Turonian-late Coniacian) of the Neuquen Group. They consist of 22 mostly articulated caudal vertebrae, a dorsal vertebra, numerous disarticulated appendicular bones and osteoderms. Autapomorphies of Mendozasaurusare: (1) subtriangular infrapostzygapophyseal fossae in anterior dorsal vertebrae; (2) postzygapostspinal laminae parallel to the plane of the postzygapophyseal facets in anterior dorsal vertebrae; (3) interzygapophyseal cavity dorsoventrally extended and limited by the spinopostzygapophyseal and spinoprezygapophyseal laminae in anterior caudal vertebrae; (4) middle caudal vertebrae slightly procoelous with reduced posterior condyles displaced dorsally; (5) laminar mid-posterior caudal neural spines with horizontal and straight dorsal border, and anterodorsal corner forming a right angle; (6) large subconic-spherical osteoderms lacking cingulum. A cladistic analysis permits the inclusion of Mendozasaurus within Titanosauridae, according to the phylogenetic definition of this clade. Although this new species exhibits almost all the titanosaurid synapomorphies proposed by some authors, it lacks prominent posterior condyles in middle caudal centra. This and other plesiomorphic traits suggest that Mendozasaurus is a basal titanosaurid, more derived than Malawisaurusin the caudal procoely. Resumen. UN NUEVO TITANOSAURIO (DINOSAURIA, SAUROPODA) DEL CRETACICO SUPERIOR DE LA PROVINCIA DE MENDOZA, ARGENTINA. Mendozasaurus neguyelap nov. gen. et sp. es un nuevo titanosaurio del Cretacico Superior de la Cuenca Neuquina y la primera especie de dinosaurio proveniente de la provincia de Mendoza, Argentina. Procede de niveles asignados preliminarmente a la Formacion Rio Neuquen (Turoniano tardio-Coniaciano tardio) del Grupo Neuquen. Esta representado por 22 vertebras caudales en su mayoria articuladas y numerosos restos desarticulados: una vertebra dorsal, arcos hemales, huesos apendiculares y grandes osteodermos. Son autapomorfias de Mendozasaurus: (1) fosas infrapostzigapofisiales subtriangulares en vertebras dorsales anteriores; (2) laminas postzigapostespinales paralelas a los planos de las facetas articulares de las postzigapofisis en vertebras dorsales anteriores; (3) cavidad interzigapofisial dorsoventralmente extendida y limitada por las laminas espinopostzigapofisial y espinoprezigapofisial en vertebras caudales anteriores; (4) centros caudales medios suavemente procelicos con condilos posteriores reducidos y desplazados dorsalmente; (5) espinas neurales de vertebras caudales medias-posteriores laminares y anteroposteriormente elongadas, con un borde dorsal horizontal y un angulo anterodorsal recto y (6) grandes osteodermos subconicos-subesfericos sin cingulum. Un analisis cladistico permite incluir a Mendozasaurusdentro de Titanosauridae, segun definiciones filogeneticas de este clado. Si bien exhibe casi todas las sinapomorfias propuestas por algunos autores para caracterizar a Titanosauridae, carece de condilos posteriores prominentes en sus vertebras caudales medias. Este y otros caracteres plesiomorficos sugieren que Mendozasaurus es un titanosaurido basal, mas derivado que Malawisaurusen su procelia caudal.