Mackinnon Memorial Hospital

Introduction. Following on from work on the European bryophyte Red List, the taxonomically and nomenclaturally updated spreadsheets used for that project have been expanded into a new checklist for the bryophytes of Europe. Methods. A steering group of ten European bryologists was convened, and over the course of a year, the spreadsheets were compared with previous European checklists, and all changes noted. Recent literature was searched extensively. A taxonomic system was agreed, and the advice and expertise of many European bryologists sought. Key results. A new European checklist of bryophytes, comprising hornworts, liverworts and mosses, is presented. Fifteen new combinations are proposed. Conclusions. This checklist provides a snapshot of the current European bryophyte flora in 2019. It will already be out-of-date on publication, and further research, particularly molecular work, can be expected to result in many more changes over the next few years.

Orkney was a major cultural center during the Neolithic, 3800 to 2500 BC. Farming flourished, permanent stone settlements and chambered tombs were constructed, and long-range contacts were sustained. From ∼3200 BC, the number, density, and extravagance of settlements increased, and new ceremonial monuments and ceramic styles, possibly originating in Orkney, spread across Britain and Ireland. By ∼2800 BC, this phenomenon was waning, although Neolithic traditions persisted to at least 2500 BC. Unlike elsewhere in Britain, there is little material evidence to suggest a Beaker presence, suggesting that Orkney may have developed along an insular trajectory during the second millennium BC. We tested this by comparing new genomic evidence from 22 Bronze Age and 3 Iron Age burials in northwest Orkney with Neolithic burials from across the archipelago. We identified signals of inward migration on a scale unsuspected from the archaeological record: As elsewhere in Bronze Age Britain, much of the population displayed significant genome-wide ancestry deriving ultimately from the Pontic-Caspian Steppe. However, uniquely in northern and central Europe, most of the male lineages were inherited from the local Neolithic. This suggests that some male descendants of Neolithic Orkney may have remained distinct well into the Bronze Age, although there are signs that this had dwindled by the Iron Age. Furthermore, although the majority of mitochondrial DNA lineages evidently arrived afresh with the Bronze Age, we also find evidence for continuity in the female line of descent from Mesolithic Britain into the Bronze Age and even to the present day.

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Abstract Bryophytes are a diverse group of organisms with unique properties, yet they are severely underrepresented in plant trait databases. Building on the recently published European Red List of bryophytes and previous trait compilations, we present the Bryophytes of Europe Traits (BET) data set, including biological traits such as those related to life history, growth habit, sexual and vegetative reproduction; ecological traits such as indicator values, substrate and habitat; and bioclimatic variables based on the species' European range. The data set includes values for 65 traits and 25 bioclimatic variables, containing more than 135,000 trait values with a completeness of 82.7% on average. The data set will enable future studies in bryophyte biology, ecology and conservation, and may help to answer fundamental questions in bryology.

The Middle Jurassic is a largely mysterious interval in dinosaur evolution, as few fossils of this age are known worldwide. In recent years, the Isle of Skye has yielded a substantial record of trackways, and a more limited inventory of body fossils, that indicate a diverse fauna of Middle Jurassic dinosaurs living in and around lagoons and deltas. Comparatively little is known about the predators in these faunas (particularly theropod dinosaurs), as their fossils are among the rarest discoveries. We here report two new isolated theropod teeth, from the Valtos Sandstone Formation and Lealt Shale Formation of Skye, which we visualized and measured using high-resolution x-ray computed microtomographic scanning (µCT) and identified via statistical and phylogenetic analyses of a large comparative dental dataset. We argue that these teeth most likely represent at least two theropod species – one small-bodied and the other large-bodied – which likely belonged to one or several clades of basal avetheropods (ceratosaurs, megalosauroids, or allosauroids). These groups, which were diversifying during the Middle Jurassic and would become dominant in the Late Jurassic, filled various niches in the food chain of Skye, probably both on land and in the lagoons. Supplementary material : Character lists, datasets, and measurements are available at https://doi.org/10.6084/m9.figshare.c.4452533

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Synopsis The first in situ dinosaur tracks from Scotland were discovered at the top of the Duntulm Formation (Bathonian, Jurassic) near to Staffin in northeastern Skye. Fifteen individual tridactyl footprints were recorded of which two pairs appear to have been part of the same trackway. The footprints are preserved as natural moulds on a mud-cracked sandstone surface. The individual track sizes range from about 30 cm to over 50 cm in length with narrow to broad digits suggestive of having been made by a medium to large bipedal dinosaur.

, but also reveals a great amount of variability within this genus, and indicates that there are currently five valid species that can be differentiated by unique combinations of dental characteristics. This variability can be included in future broad-scale cladistics analyses of atoposaurids and closely related crocodyliforms, which promise to help untangle the complicated taxonomy and evolutionary history of Atoposauridae. © 2015 The Authors. Zoological Journal of the Linnean Society published by John Wiley & Sons Ltd on behalf of The Linnean Society of London.

Tracks of a juvenile theropod dinosaur with footprint lengths of between 2 and 9 cm as well as adults of the same ichnospecies with footprints of about 15–25 cm in length were found in the Bathonian (Middle Jurassic) Kilmaluag Formation of Score Bay, northwestern Trotternish Peninsula, Isle of Skye, Scotland, UK. Two footprint sizes occur together on the same bedding plane in the central portion of Score Bay, both in situ and on loose blocks. Another horizon containing footprints above this was also identified. The footprints from the lowest horizon were produced in a desiccated silty mud that was covered with sand. A close association of both adults and juveniles with similar travel direction indicated by the footprints may suggest post-hatching care in theropod dinosaurs. Other footprints, produced on a rippled sandy substrate, have been found on the slightly higher bedding plane at this locality. Loose blocks found 130 m to the northeast in the central part of Score Bay have not been correlated with any in situ sediments, but were preserved in a similar manner to those from the higher bedding plane. These tracks represent the youngest dinosaur remains yet found in Scotland.

Introduction. Many taxa have been added to the bryophyte flora of Britain and Ireland since the publication of the previous checklist in 2008. This has coincided with numerous taxonomic and nomenclatural changes in the European flora, especially as a consequence of molecular sequencing. With the publication of a new European checklist in 2020 it is now appropriate to provide an updated and annotated list for Britain and Ireland.Methods. A working group was convened to consider and reach a consensus on taxonomic and nomenclatural matters arising from the European checklist. Special consideration was given to the citation of the authors of names. Taxa reported as new to Britain and Ireland since the previous checklist in 2008 were incorporated.Key results. A new checklist of bryophytes (hornworts, liverworts and mosses) is presented. Based on the taxonomy adopted in the checklist, the bryophyte flora of Britain and Ireland comprises four hornworts (Anthocerotophyta), 299 liverworts (Marchantiophyta), and 795 mosses (Bryophyta) at species rank, with 42 additional infraspecific taxa.

New national and regional bryophyte records, This is the first record of Campylopus subulatus, Racomitrium elongatum and Taxiphyllum wissgrillii for the Liguria Region.
\nCampylopus subulatus was found on small serpentinite outcrops occupied by a discontinuous plant community, with a paucispecific contingent of phanerogams dominated by succulent chamaephytes and therophytes (refered to the Sedo-Scleranthetalia order) and a richer contingent of bryophytes including Racomitrium elongatum Frisvoll, Hypnum callichroum Brid., Ceratodon purpureus (Hedw.) Brid., Grimmia donniana Sm., Dicranum scoparium Hedw. and Bryum Hedw. spp. These outcrops occur in a marshy area, mainly characterised by mountain grasslands and beech forests growing in a sub-oceanic climate. Racomitrium elongatum was found on serpentinite outcrops within the Beigua Regional Natural Park, in a beech forest and in marshy mountain grassland. Taxiphyllum wissgrillii was found a serpentinite outcrop in a deciduous mesophilic forest dominated by beech (Fagus sylvatica L., referring to the Fagion silvaticae alliance), in the Beigua Regional Natural Park.

The Middle Jurassic was a dynamic interval in dinosaur evolution, but the dinosaur fossil record from this time is extremely poor throughout the world. The Isle of Skye (Scotland) preserves marginal marine and terrestrial deposits of Middle Jurassic age, which have yielded sparse bones, teeth, footprints and small segments of trackways belonging to dinosaurs. We report the discovery of the most extensive dinosaur fossil site yet known in Scotland: a coastal outcrop of the Duntulm Formation (Bathonian) at Cairidh Ghlumaig, Skye that preserves numerous trackways of sauropod dinosaurs in multiple layers deposited in a lagoonal system. We present an initial description of these tracks and identify them as most likely belonging to a primitive, non-neosauropod species that retained a large claw on manual digit I and produced narrow-gauge trackways. They provide additional evidence that basal sauropods persisted deep into the Middle Jurassic, a time when the earliest members of larger and more derived sauropod lineages were radiating. The new Skye tracks document multiple generations of sauropods living within the lagoonal environments of Jurassic Scotland, and along with other tracks found over the past two decades, suggest that sauropods may have frequented such environments, contrary to their image as land-bound behemoths.

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Globally there is increasing concern about the decline in the health of oak Quercus trees. The impact of a decline in oak trees on associated biodiversity, species that utilize oak trees, is unknown. Here we collate a database of all known birds, bryophytes, fungi, invertebrates, lichens and mammals that use oak (Quercus petraea and Q. robur) in the UK. In total 2300 species are listed in the database. For each species we provide a level of association with oak, ranging from obligate (only found on oak) to cosmopolitan (found on a wide range of other tree species). Data on the ecology of each oak associated species was collated: part of tree used, use made of tree (feeding, roosting, breeding), age of tree, woodland type, tree form (coppice, pollarded, or natural growth form) and season when the tree was used. Data on use or otherwise by each of the 2300 species of 30 other tree species was also collated. A complete list of data sources is provided. For further insights into how this data can be used see Collapsing foundations: The ecology of the British oak, implications of its decline and mitigation options [1]. Data can be found at EIDC https://doi.org/10.5285/22b3d41e-7c35-4c51-9e55-0f47bb845202.

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Fossils of Mesozoic vertebrates are rare in Scotland, particularly specimens of marine reptiles such as plesiosaurs and ichthyosaurs. We describe a suite of ichthyosaur fossils from the Early to Middle Jurassic of Skye, which to our knowledge are the first ichthyosaurs from Scotland to be described and figured in detail. These fossils span approximately 30 million years, from the Sinemurian to the Bathonian, and indicate that ichthyosaurs were a major component of Scottish marine faunas during this time. The specimens include isolated teeth that could represent the most northerly known occurrences of the widespread Sinemurian species Ichthyosaurus communis , a characteristic component of the famous Lyme Regis faunas of England, suggesting that such faunas were also present in Scotland during the Early Jurassic. An associated humerus and vertebrae from Toarcian–Bajocian-aged deposits are named as a new genus and species of basal neoichthyosaurian, Dearcmhara shawcrossi . The taxonomic affinities of this taxon, which comes from a critical but poorly sampled interval in the fossil record, suggest that non-ophthalmosaurid neoichthyosaurians dominated European assemblages around the Early–Middle Jurassic boundary, and were later replaced by ophthalmosaurids, whose radiation likely took place outside Europe. Many of these specimens were collected by amateurs and donated to museum collections, a co-operative relationship essential to the preservation of Scotland’s fossil heritage.

ABSTRACT The Middle Jurassic is a poorly sampled time interval for non-pelagic neosuchian crocodyliforms, which obscures our understanding of the origin and early evolution of major clades. Here we report a lower jaw from the Middle Jurassic (Bathonian) Duntulm Formation of the Isle of Skye, Scotland, UK, which consists of an isolated and incomplete left dentary and part of the splenial. Morphologically, the Skye specimen closely resembles the Cretaceous neosuchians Pachycheilosuchus and Pietraroiasuchus , in having a proportionally short mandibular symphysis, shallow dentary alveoli and inferred weakly heterodont dentition. It differs from other crocodyliforms in that the Meckelian canal is dorsoventrally expanded posterior to the mandibular symphysis and drastically constricted at the 7th alveolus. The new specimen, together with the presence of Theriosuchus sp. from the Valtos Formation and indeterminate neosuchians from the Kilmaluag Formation, indicates the presence of a previously unrecognised, diverse crocodyliform fauna in the Middle Jurassic of Skye, and Europe more generally. Small-bodied neosuchians were present, and ecologically and taxonomically diverse, in nearshore environments in the Middle Jurassic of the UK.

1. Including the present case report, nine instances of open dislocation of the elbow have been encountered since the turn of the century. In all except the present instance there was disruption of the brachial vessels at the antecubital fossa while the median nerve remained intact. 2. Five cases of closed dislocation of the elbow with disruption of the brachial vessels have been observed since 1900. Immediate surgical intervention is needed in this group as well as in the open dislocations. 3. It is not only unnecessary but undesirable to attempt to re-establish the continuity of the brachial vessels at the antecubital fossa because of the rich collateral circulation that is known to exist.

Middle Jurassic dinosaur fossils are exceedingly rare, but new discoveries from the Isle of Skye, Scotland, are beginning to fill this gap. We here describe a new dinosaur tracksite found in the Lealt Shale Formation (Bathonian) of the Great Estuarine Group at Rubha nam Brathairean (Brothers' Point) on Skye. The site preserves an abundance of small sauropod manus and pes prints and several isolated and broken medium-to-large tridactyl footprints. The main site occurs on a single horizon of shaley limestone that formed in a lagoonal environment. The sauropod tracks are tentatively assigned to the ichnotaxon Breviparopus due to the narrow gauge of the trackways, the digital characteristics of the pes, and the ratio of heteropody observed between the manus and the pes. A theropod trackmaker is inferred for some of the tridactyl impressions with several indicative of the ichnotaxon Eubrontes . This new site strengthens the inference, originally based on a previously discovered locality near Duntulm Castle (Duntulm Formation) in northern Skye, that sauropods habitually spent time in lagoons during the Middle Jurassic. Supplementary material : The photogrammetric model of track BP2_40 and associated metadata and photographs are available at https://doi.org/10.6084/m9.figshare.c.4046390